Temporal changes of taxonomic and functional diversity in dung beetles inhabiting forest fragments and pastures in Los Tuxtlas Biosphere Reserve, Mexico

We evaluated short- (among months within-years) and long-term (between 1999-2000 2016-2017) temporal patterns of taxonomic functional β- diversity (and its components substitution gain/loss) dung beetle assemblages in forest fragments pastures the Los Tuxtlas Biosphere Reserve (LTBR). Habitat type affected dissimilarity richness difference component, with average values being respectively 1.42 1.56 times higher than fragments. Only habitat was important for richness, 1.93 1.69 2016-2017, respectively. Pastures were taxonomically functionally poorer but also more temporally dynamic both within-year between-years. is a determining factor dynamics, presenting stable cattle pastures.